

During a field trip, Shio is revealed to be an "E-Gene Holders", a person who wield weapons infused with the spirits of historical figures. The soul of Oda Nobunaga awakens after she tries to rescue a friend.
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Nobunaga Shimazaki
Mahesh Mirza (voice)

Nobuo Tobita
Nobunaga Oda (voice)

Ayumu Murase
Gaudi Antoni (voice)

Akeno Watanabe
Dogoo (voice)

Sumire Uesaka
Galileo Galilei (voice)

Chiwa Saito
Geronimo (voice)

Yuu Asakawa
Isaac Newton (voice)

Makoto Yasumura
John Hunter (voice)

Koji Yusa
Saint-Germain (voice)

Ai Satou
Shirei (voice)

Kenjiro Tsuda
Robert Capa (voice)

Tatsuhisa Suzuki
Jack Kirisaki (voice)

Nobunaga Shimazaki
Gandhi Mahatma (voice)

Akira Ishida
Francois Vidocq (voice)
1 season available

Weapon otaku Ogura Sio is on a school trip to Taiwan when she and her classmates get caught up in an Evolutionary Invasion Object attack, not that she has any idea what is going on. And who is the guy with the knives?

The incident in Taiwan draws to a close, and Sio gets a chance to have a heart-to-heart with recovering Asao before deciding whether she will join the monster-fighting team or not.

Sio starts intensive training at the Dogoo facility with war photographer Robert Capa. She manages to make some friends on base, but the reality of everyone's situation there is harsh.

Ogura and the others are sent to deal with an Evolutionary Invasion Object that can ride a hurricane.

Sio is assigned to the 2nd platoon. When she joins up with them on their airship, she learns that e-gene holders just keep getting weirder. Meanwhile, DOGOO learns something new about the Evolutionary Invaders--and it's not good news.

Gaudi, Cyx, and Geronimo of the First Platoon crash-land on a snowy mountain. By coincidence, they come across a research outpost where something has gone horribly wrong...

When a famed sunken battleship of the Japanese Imperial Navy starts wreaking havoc near the Panama Canal, Domoo realizes it's due to Evolutionary Invasion Objects, and sends in the Second Platoon

Vidocq discovers something important about how the Evolutionary Invaders are communicating. It's time for the 1st platoon to move out--and when the mission gets a little crazy, the 2nd platoon is called in to back them up.

Evolutionary Invasion Objects suddenly show up during Kaoru's visit to Sio, who is convalescing in a hospital in Japan.

As the plan to deal with the Objects in the tunnel takes shape, the commander remembers the ups and downs of her extra-long life.

Will Nobunagun's plan to capture the Evolutionary Invasion Object tunnel and destroy the original cells work? Nine E-gene holders begin Dogoo's most important operation yet.

Jack and Geronimo make a bet; Hunter has to learn to fight however he can. Meanwhile, Nobunagun faces a new threat.

Who is Jack the Ripper? Sio gets let in on the secret as Operation Stone Forest comes to an end. And then?